🌸

Flower Development and ABC Model

Sep 1, 2025

Overview

This lecture explains the ABC model of flowering, focusing on gene classes, gene combinations, and their effects on flower organ identity and mutations.

Structure of a Flower

  • A typical flower has four main components: sepal, petal, stamen, and carpel.
  • These components are arranged from outermost to innermost: sepal, petal, stamen, carpel ("sapestica" mnemonic).

ABC Model Gene Classes

  • Three classes of genes regulate flower development: Class A, Class B, and Class C.
  • Class A and Class C genes have antagonistic (opposing) relationships; if one is missing, the other takes over.
  • Each gene class has specific gene examples:
    • Class A: APETALA1 (AP1), APETALA2 (AP2)
    • Class B: APETALA3 (AP3), PISTILLATA (PI)
    • Class C: AGAMOUS (AG)

Combination of Gene Expression and Organ Identity

  • Flower organs develop based on combinations of gene class expression across four "whorls" (concentric regions).
  • Whorl 1: Class A genes alone → Sepal forms.
  • Whorl 2: Class A + Class B genes → Petal forms.
  • Whorl 3: Class B + Class C genes → Stamen forms.
  • Whorl 4: Class C genes alone → Carpel forms.

Gene Mutations and Effects

  • Class A Mutation: Class C replaces A, resulting in flowers with only carpels and stamens.
  • Class C Mutation: Class A replaces C, resulting in flowers with only sepals and petals.
  • Class B Mutation: Only Class A and C are present, leading to flowers with only sepals and carpels.

Key Terms & Definitions

  • Whorl — A concentric ring in a flower where specific organs form.
  • Class A gene — Gene group defining sepals and petals (e.g., AP1, AP2).
  • Class B gene — Gene group defining petals and stamens (e.g., AP3, PI).
  • Class C gene — Gene group defining stamens and carpels (e.g., AG).
  • Antagonistic genes — Genes whose activities oppose each other (here, Class A and C).

Action Items / Next Steps

  • Review the roles of AP1/AP2, AP3/PI, and AG genes in flower organ identity.
  • Practice drawing and labeling the ABC model triangle and predicting outcomes of gene mutations.