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Photosystems in Photosynthesis

Jun 16, 2025

Overview

This lecture covers photosystems in photosynthesis, focusing on their structure, function, and the role of pigments in absorbing light energy.

Photosystem Structure

  • Photosystems are complexes found in the thylakoid membrane involved in photosynthesis.
  • Each photosystem contains multiple pigment molecules organized into accessory pigments and a reaction center.
  • The reaction center of each photosystem contains two chlorophyll a molecules.

Types of Pigments

  • Accessory pigments absorb light of various wavelengths and pass the energy to the reaction center.
  • Different pigments maximize light absorption by capturing a spectrum of wavelengths (400–700 nm).
  • The reaction center is the site where absorbed light energy is funneled and utilized.

Function of Photosystems

  • Photosystems absorb light via pigments, transferring energy to the reaction center.
  • Upon receiving enough energy, the reaction center becomes photoactivated and releases a high-energy electron.
  • The primary role is to absorb light, activate the reaction center, and initiate electron release.

Photosystem I vs. Photosystem II

  • Both photosystems have similar structure and function regarding light absorption and electron release.
  • Photosystem II uniquely contains the oxygen-evolving complex enzyme, which participates in a later step.

Electron Replacement in the Reaction Center

  • When the reaction center releases an electron, it must be replaced to maintain structural stability.
  • Continuous loss of electrons without replacement would destabilize the reaction center.

Key Terms & Definitions

  • Photosystem — Protein complex in thylakoids that absorbs light and releases electrons during photosynthesis.
  • Accessory Pigments — Pigments that absorb different wavelengths of light and transfer energy to chlorophyll a.
  • Reaction Center — The core of the photosystem, containing two chlorophyll a molecules, where photoactivation occurs.
  • Photoactivation — Process where the reaction center absorbs enough energy to release a high-energy electron.
  • Oxygen Evolving Complex — Enzyme present in photosystem II involved in water splitting (not detailed here).

Action Items / Next Steps

  • Review the structure and function of photosystems, focusing on pigment roles and electron flow.
  • Prepare for detailed study of the light-dependent reactions in upcoming lessons.