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Understanding Deuterostomes and Their Evolution

Sep 16, 2024

Phylogenetic Tree: Deuterostomes

Overview

  • The lecture focuses on deuterostomes, particularly echinoderms and basal chordates.
  • Distinction between protostomes and deuterostomes:
    • Protostomes: First invagination in the embryo develops into the mouth.
    • Deuterostomes: First invagination becomes the anus; the mouth develops from the second invagination.
  • Importance of studying echinoderms for understanding chordate development.

Echinoderms

  • Examples: Starfish, sea urchins, and sea cucumbers.
  • Sea Cucumber: Elongated body; mouth and anus at opposite ends.
  • Sea Urchin: Ventral structure.
  • Starfish: Often mistaken for having five-fold symmetry but actually exhibit bilateral symmetry.

Chordates

  • Sister group to echinoderms.
  • Includes basal chordates and vertebrates.

Basal Chordates

  • Tunicates: Start life as typical chordates but later embed into substrates, resembling sponges.
  • Amphioxus (Cephalochordate): Small organisms with a head region.

Craniata

  • Includes hagfish and vertebrates.
  • Hagfish (Hyperartia):
    • Lack lower jaw and limbs.
    • Possess slime glands for defense (similar to the character "Slimer" from Ghostbusters).
  • Lampreys:
    • Have eyes and oral discs with teeth.
    • Evolving fins for improved swimming.

Nathostomata

  • Vertebrates with jaws (biting mouths).
  • Basal Group: Chondrichthyes
    • Includes cartilaginous fish like sharks and rays.
    • Lack bones; strong supportive structures are unnecessary in aquatic environments.

Fish with Bone

  • Ray-finned Fish (Actinopterygii):
    • Common fish such as tuna and salmon.
    • Includes seahorses.
  • Lobe-finned Fish (Sarcopterygii):
    • Ancestors to terrestrial vertebrates.
    • Includes lungfish and terrestrial vertebrates (mammals, reptiles, birds, amphibians).

Lungfish (Dipnoi)

  • Three types based on geography: Australia, South America, Africa.
  • Have muscular limbs; some capable of terrestrial movement.
  • Possess bone in fins, unlike ray-finned fish.

Evolution to Terrestrial Vertebrates

  • Terrestrial vertebrates evolved from sarcopterygian ancestors.
  • Example: Fossil fish from the Devonian period.
  • Bone structure in fins:
    • Single large bone connects to the body.
    • Pair of bones, followed by smaller bones and five digits.
    • This bone pattern is inherited by modern terrestrial vertebrates.

Conclusion

  • Evolutionary homology in limb structure can be traced back to these sarcopterygian ancestors, illustrating the continuity from aquatic to terrestrial life forms.