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Light-Dependent Photosynthesis Overview

Jul 19, 2025

Overview

This lecture covers the light-dependent stage of photosynthesis, focusing on the photolysis of water, ATP production, and the formation of reduced NADP.

Structure and Location of the Light-Dependent Stage

  • The light-dependent stage is the first part of photosynthesis and occurs on the thylakoid membranes inside the grana of chloroplasts.
  • Thylakoid membranes are organized into stacks called grana.

Steps of the Light-Dependent Stage

  • The process is divided into three main steps: photolysis of water, production of ATP (photophosphorylation), and formation of reduced NADP.

Photolysis of Water

  • Photolysis splits one water molecule (H₂O) into oxygen (O₂), electrons, and hydrogen ions (protons) using sunlight energy.
  • The reaction is catalyzed by an enzyme within photosystem II (PSII) on the thylakoid membrane.
  • From each water molecule, two electrons and two hydrogen ions are produced.
  • The oxygen formed is used by the plant for respiration or diffuses out through stomata if not needed.
  • The electrons and protons generated are essential for the next step, photophosphorylation.

Production of ATP (Photophosphorylation)

  • ADP (adenosine diphosphate) combines with an inorganic phosphate group in the presence of light to form ATP (adenosine triphosphate).
  • This process is called photophosphorylation: "photo" for light, "phosphorylation" for adding a phosphate group.

Formation of Reduced NADP

  • NADP (nicotinamide adenine dinucleotide phosphate) acts as a coenzyme and accepts hydrogen atoms, becoming reduced NADP.
  • Reduced NADP is produced at the end of the light-dependent reactions.

Key Terms & Definitions

  • Thylakoid membrane — site within chloroplasts where the light-dependent reactions occur.
  • Grana (Granum) — stacks of thylakoid membranes.
  • Photolysis — splitting of water using light energy.
  • Photosystem II (PSII) — protein complex involved in capturing light and catalyzing photolysis.
  • Photophosphorylation — production of ATP from ADP and phosphate using light energy.
  • NADP — coenzyme that accepts hydrogen to become reduced NADP.
  • Stomata — pores in leaves for gas exchange.

Action Items / Next Steps

  • Review the steps and key terms of the light-dependent stage.
  • Prepare for further study on the next stages of photosynthesis.